Problems in discrimination of tridactyl dinosaur footprints, exemplified by the Hettangian trackways, the causses, France

Abstract

Numerous tridactyl dinosaur footprints have been discovered in the Lower Jurassic (Hettangian) of the Causses, France. They exhibit the usual Grallator‐like morphology in which differences seem to be more a result of normal intraspecific variability than of inherent differences between skeletons. However, such subjective conclusions beg an analytical confirmation, which was the principal objective of this study. For more reliable shape determinations, typical specimens from well‐preserved footprint ichnofaunas of the Connecticut Valley (USA) were used. Statistical methods are necessary to verify the homogeneity of such footprint populations. Determinations of the variability, confidence interval for the mean, and ratios between length characters are particularly important in reducing the influence of the size of each print. By this method, the ichnospecies Anomoepus intermedius, Eubrontes giganteus, and Grallator sillimani are statistically distinct; the equally distinct new ichnospecies from Saint‐Léons (Aveyron) can be designated as Grallator lescurei and the trackways of Saint‐Laurent as G. minusculus. A problem remains in classifying the trackmaking dinosaurs. Their prior assignment to the theropods apparently should be revised because of the abundance of grallatorid footprints in the Lower Jurassic, which seems to contradict conclusions drawn from paleoecological data.