Gradient and Habitat Continuum era
Henry Gleason championed the individualistic concept of plant associations, arguing that species distributions reflect continuous environmental gradients rather than tight community integration. Victor Shelford helped institutionalize ecogeography and habitat-based reasoning, linking climate, soils, and geography to the distribution of plant and animal communities. Carl Troll pioneered landscape ecology and gradient-based climate-vegetation frameworks, organizing vegetation into gradient zones and eco-regions to predict turnover. Arthur Tansley introduced the ecosystem concept, integrating the physical environment, vegetation, and biotic interactions into a unit that underpins habitat-continuum biogeography.