A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China

Abstract

The Southeast Asian xyleborine ambrosia beetle fauna is reviewed for the first time. Thirty-four genera and 315 species are reviewed, illustrated, and keyed to genera and species. Sixty-three new species are described: <i>Amasa cycloxyster</i> <b>sp. nov.</b>, <i>Amasa galeoderma</i> <b>sp. nov.</b>, <i>Amasa gibbosa</i> <b>sp. nov.</b>, <i>Amasa lini</i> <b>sp. nov.</b>, <i>Amasa tropidacron</i> <b>sp. nov.</b>, <i>Amasa youlii</i> <b>sp. nov.</b>, <i>Ambrosiophilus caliginestris</i> <b>sp. nov.</b>, <i>Ambrosiophilus indicus</i> <b>sp. nov.</b>, <i>Ambrosiophilus lannaensis</i> <b>sp. nov.</b>, <i>Ambrosiophilus papilliferus</i> <b>sp. nov.</b>, <i>Ambrosiophilus wantaneeae</i> <b>sp. nov.</b>, <i>Anisandrus achaete</i> <b>sp. nov.</b>, <i>Anisandrus auco</i> <b>sp. nov.</b>, <i>Anisandrus auratipilus</i> <b>sp. nov.</b>, <i>Anisandrus congruens</i> <b>sp. nov.</b>, <i>Anisandrus cryphaloides</i> <b>sp. nov.</b>, <i>Anisandrus feronia</i> <b>sp. nov.</b>, <i>Anisandrus hera</i> <b>sp. nov.</b>, <i>Anisandrus paragogus</i> <b>sp. nov.</b>, <i>Anisandrus sinivali</i> <b>sp. nov.</b>, <i>Anisandrus venustus</i> <b>sp. nov.</b>, <i>Anisandrus xuannu</i> <b>sp. nov.</b>, <i>Arixyleborus crassior</i> <b>sp. nov.</b>, <i>Arixyleborus phiaoacensis</i> <b>sp. nov.</b>, <i>Arixyleborus setosus</i> <b>sp. nov.</b>, <i>Arixyleborus silvanus</i> <b>sp. nov.</b>, <i>Arixyleborus sittichayai</i> <b>sp. nov.</b>, <i>Arixyleborus titanus</i> <b>sp. nov.</b>, <i>Coptodryas amydra</i> <b>sp. nov.</b>, <i>Coptodryas carinata</i> <b>sp. nov.</b>, <i>Coptodryas inornata</i> <b>sp. nov.</b>, <i>Cyclorhipidion amasoides</i> <b>sp. nov.</b>, <i>Cyclorhipidion amputatum</i> <b>sp. nov.</b>, <i>Cyclorhipidion denticauda</i> <b>sp. nov.</b>, <i>Cyclorhipidion muticum</i> <b>sp. nov.</b>, <i>Cyclorhipidion obesulum</i> <b>sp. nov.</b>, <i>Cyclorhipidion petrosum</i> <b>sp. nov.</b>, <i>Cyclorhipidion truncaudinum</i> <b>sp. nov.</b>, <i>Cyclorhipidion xeniolum</i> <b>sp. nov.</b>, <i>Euwallacea geminus</i> <b>sp. nov.</b>, <i>Euwallacea neptis</i> <b>sp. nov.</b>, <i>Euwallacea subalpinus</i> <b>sp. nov.</b>, <i>Euwallacea testudinatus</i> <b>sp. nov.</b>, <i>Heteroborips fastigatus</i> <b>sp. nov.</b>, <i>Heteroborips indicus</i> <b>sp. nov.</b>, <i>Microperus latesalebrinus</i> <b>sp. nov.</b>, <i>Microperus minax</i> <b>sp. nov.</b>, <i>Microperus sagmatus</i> <b>sp. nov.</b>, <i>Streptocranus petilus</i> <b>sp. nov.</b>, <i>Truncaudum bullatum</i> <b>sp. nov.</b>, <i>Xyleborinus cuneatus</i> <b>sp. nov.</b>, <i>Xyleborinus disgregus</i> <b>sp. nov.</b>, <i>Xyleborinus echinopterus</i> <b>sp. nov.</b>, <i>Xyleborinus ephialtodes</i> <b>sp. nov.</b>, <i>Xyleborinus huifenyinae</i> <b>sp. nov.</b>, <i>Xyleborinus jianghuansuni</i> <b>sp. nov.</b>, <i>Xyleborinus thaiphami</i> <b>sp. nov.</b>, <i>Xyleborinus tritus</i> <b>sp. nov.</b>, <i>Xyleborus opacus</i> <b>sp. nov.</b>, <i>Xyleborus sunisae</i> <b>sp. nov.</b>, <i>Xyleborus yunnanensis</i> <b>sp. nov.</b>, <i>Xylosandrus bellinsulanus</i> <b>sp. nov.</b>, <i>Xylosandrus spinifer</i> <b>sp. nov.</b>. Thirteen new combinations are given: <i>Ambrosiophilus consimilis</i> (Eggers) <b>comb. nov.</b>, <i>Anisandrus carinensis</i> (Eggers) <b>comb. nov.</b>, <i>Anisandrus cristatus</i> (Hagedorn) <b>comb. nov.</b>, <i>Anisandrus klapperichi</i> (Schedl) <b>comb. nov.</b>, <i>Anisandrus percristatus</i> (Eggers) <b>comb. nov.</b>, <i>Arixyleborus resecans</i> (Eggers) <b>comb. nov.</b>, <i>Cyclorhipidion armiger</i> (Schedl) <b>comb. nov.</b>, <i>Debus quadrispinus</i> (Motschulsky) <b>comb. nov.</b>, <i>Heteroborips tristis</i> (Eggers) <b>comb. nov.</b>, <i>Leptoxyleborus machili</i> (Niisima) <b>comb. nov.</b>, <i>Microperus cruralis</i> (Schedl) <b>comb. nov.</b>, <i>Planiculus shiva</i> (Maiti & Saha) <b>comb. nov.</b>, <i>Xylosandrus formosae</i> (Wood) <b>comb. nov.</b> Twenty-four new synonyms are proposed: <i>Ambrosiophilus osumiensis</i> (Murayama, 1934) (= <i>Xyleborus nodulosus</i> Eggers, 1941 <b>syn. nov.</b>); <i>Ambrosiophilus subnepotulus</i> (Eggers, 1930) (= <i>Xyleborus cristatuloides</i> Schedl, 1971 <b>syn. nov.</b>); <i>Ambrosiophilus sulcatus</i> (Eggers, 1930) (= <i>Xyleborus sinensis</i> Eggers, 1941 <b>syn. nov.</b>; = <i>Xyleborus sulcatulus</i> Eggers, 1939 <b>syn. nov.</b>); <i>Anisandrus hirtus</i> (Hagedorn, 1904) (= <i>Xyleborus hirtipes</i> Schedl, 1969 <b>syn. nov.</b>); <i>Cnestus protensus</i> (Eggers, 1930) (= <i>Cnestus rostratus</i> Schedl, 1977 <b>syn. nov.</b>); <i>Cyclorhipidion bodoanum</i> (Reitter, 1913) (= <i>Xyleborus misatoensis</i> Nobuchi, 1981 <b>syn. nov.</b>); <i>Cyclorhipidion distinguendum</i> (Eggers, 1930) (= <i>Xyleborus fukiensis</i> Eggers, 1941 <b>syn. nov.</b>; = <i>Xyleborus ganshoensis</i> Murayama, 1952 <b>syn. nov.</b>); <i>Cyclorhipidion inarmatum</i> (Eggers, 1923) (= <i>Xyleborus vagans</i> Schedl, 1977 <b>syn. nov.</b>); <i>Debus quadrispinus</i> (Motschulsky, 1863) (= <i>Xyleborus fallax</i> Eichhoff, 1878 <b>syn. nov.</b>); <i>Euwallacea gravelyi</i> (Wichmann, 1914) (= <i>Xyleborus barbatomorphus</i> Schedl, 1951 <b>syn. nov.</b>); <i>Euwallacea perbrevis</i> (Schedl, 1951) (= <i>Xyleborus molestulus</i> Wood, 1975 <b>syn. nov.</b>; <i>Euwallacea semirudis</i> (Blandford, 1896) (= <i>Xyleborus neohybridus</i> Schedl, 1942 <b>syn. nov.</b>); <i>Euwallacea sibsagaricus</i> (Eggers, 1930) (= <i>Xyleborus tonkinensis</i> Schedl, 1934 <b>syn. nov.</b>); <i>Euwallacea velatus</i> (Sampson, 1913) (= <i>Xyleborus rudis</i> Eggers, 1930 <b>syn. nov.</b>); <i>Microperus kadoyamaensis</i> (Murayama, 1934) (= <i>Xyleborus pubipennis</i> Schedl, 1974 <b>syn. nov.</b>; =<i>Xyleborus denseseriatus</i> Eggers, 1941 <b>syn. nov.</b>); <i>Stictodex dimidiatus</i> (Eggers, 1927) (=<i>Xyleborus dorsosulcatus</i> Beeson, 1930 <b>syn. nov.</b>); <i>Webbia trigintispinata</i> Sampson, 1922 (= <i>Webbia mucronatus</i> Eggers, 1927 <b>syn. nov.</b>); <i>Xyleborinus artestriatus</i> (Eichhoff, 1878) (= <i>Xyelborus angustior</i> [<i>sic</i>] Eggers, 1925 <b>syn. nov.</b>; = <i>Xyleborus undatus</i> Schedl, 1974 <b>syn. nov.</b>); <i>Xyleborinus exiguus</i> (Walker, 1859) (= <i>Xyleborus diversus</i> Schedl, 1954 <b>syn. nov.</b>); <i>Xyleborus muticus</i> Blandford, 1894 (= <i>Xyleborus conditus</i> Schedl, 1971 <b>syn. nov.</b>; = <i>Xyleborus lignographus</i> Schedl, 1953 <b>syn. nov.</b>). Seven species are removed from synonymy and reinstated as valid species: <i>Anisandrus cristatus</i> (Hagedorn, 1908), <i>Cyclorhipidion tenuigraphum</i> (Schedl, 1953), <i>Diuncus ciliatoformis</i> (Schedl, 1953), <i>Euwallacea gravelyi</i> (Wichmann, 1914), <i>Euwallacea semirudis</i> (Blandford, 1896), <i>Microperus fulvulus</i> (Schedl, 1942), <i>Xyleborinus subspinosus</i> (Eggers, 1930).