Catalogue of Tenebrionidae (Coleoptera) of North America

Abstract

This catalogue includes all valid family-group (8 subfamilies, 52 tribes, 14 subtribes), genus-group (349 genera, 86 subgenera), and species-group names (2825 species, 215 subspecies) of darkling beetles (Coleoptera: Tenebrionidae) known to occur in North America and their available synonyms. Data on extant, subfossil and fossil taxa are given. For each name the author and year and page number of the description are provided, with additional information (e.g., type species for genus-group names, author of synonymies for invalid taxa) depending on the taxon rank. Several new nomenclatural acts are included. One new genus, <i>Lepidocnemeplatia</i> Bousquet and Bouchard, is described. <i>Spelaebiosis</i> Bousquet and Bouchard [for <i>Ardoinia</i> Özdikmen, 2004], <i>Blapstinus marcuzzii</i> Aalbu [for <i>Blapstinus kulzeri</i> Marcuzzi, 1977], and <i>Hymenorus campbelli</i> Bouchard [for <i>Hymenorus oculatus</i> Doyen and Poinar, 1994] are proposed as new replacement names. Supporting evidence is provided for the conservation of usage of <i>Tarpela micans</i> (Fabricius, 1798) <i>nomen protectum</i> over <i>Tarpela vittata</i> (Olivier, 1793) <i>nomen oblitum</i>. The generic names <i>Psilomera</i> Motschulsky, 1870 [= <i>Stenomorpha</i> Solier, 1836], <i>Steneleodes</i> Blaisdell, 1909 [= <i>Xysta</i> Eschscholtz, 1829], <i>Ooconibius</i> Casey, 1895 and <i>Euconibius</i> Casey, 1895 [= <i>Conibius</i> LeConte, 1851] are new synonyms (valid names in square brackets). The following 127 new synonymies of species-group names, listed in their original combination, are proposed (valid names, in their current combination, placed in square brackets): <i>Bothrasida mucorea</i> Wilke, 1922 [= <i>Pelecyphorus guanajuatensis</i> (Champion, 1884)]; <i>Parasida zacualpanicola</i> Wilke, 1922 [= <i>Pelecyphorus asidoides</i> Solier, 1836]; <i>Stenosides kulzeri</i> Pallister, 1954, <i>Stenosides bisinuatus</i> Pallister, 1954, and <i>Parasida trisinuata</i> Pallister, 1954 [= <i>Pelecyphorus dispar</i> (Champion, 1892)]; <i>Asida favosa</i> Champion, 1884 and <i>Asida similata</i> Champion, 1884 [= <i>Pelecyphorus fallax</i> (Champion, 1884)]; <i>Ologlyptus bicarinatus</i> Champion, 1884 [= <i>Pelecyphorus indutus</i> (Champion, 1884)]; <i>Parasida laciniata</i> Casey, 1912 and <i>Parasida cristata</i> Pallister, 1954 [= <i>Pelecyphorus liratus</i> (LeConte, 1854)]; <i>Parasida esperanzae</i> Wilke, 1922 and <i>Parasida mixtecae</i> Wilke, 1922 [= <i>Pelecyphorus longipennis</i> (Champion, 1884)]; <i>Parasida tolucana</i> Casey, 1912 [= <i>Pelecyphorus scutellaris</i> (Champion, 1884)]; <i>Parasida purpusi</i> Wilke, 1922 [= <i>Pelecyphorus tristis</i> (Champion, 1884)]; <i>Astrotus nosodermoides</i> Champion, 1892 [= <i>Pelecyphorus erosus</i> (Champion, 1892)]; Astrotus seticornis var. humeralis Champion, 1884 [= <i>Pelecyphorus seticornis</i> (Champion, 1884)]; <i>Pactostoma breviuscula</i> Casey, 1912, <i>Pactostoma exoleta</i> Casey, 1912, <i>Pactostoma luteotecta</i> Casey, 1912, <i>Pactostoma monticola</i> Casey, 1912, <i>Pactostoma obtecta</i> Casey, 1912, and <i>Pactostoma sigillata</i> Casey, 1912 [=<i>Pelecyphorus anastomosis</i> (Say, 1824)]; <i>Ologlyptus canus</i> Champion, 1884 and <i>Ologlyptus sinuaticollis</i> Champion, 1884 [= <i>Pelecyphorus graciliformis</i> (Solier, 1836)]; <i>Gonasida elata reducta</i> Casey, 1912, <i>Gonasida elata prolixa</i> Casey, 1912, and <i>Gonasida aucta</i> Casey, 1912 [= <i>Philolithus elatus compar</i> (Casey, 1912)]; <i>Gonasida alaticollis</i> Casey, 1912 [= <i>Philolithus elatus difformis</i> (LeConte, 1854)]; <i>Gonasida gravida</i> Casey, 1912 [= <i>Philolithus elatus elatus</i> (LeConte, 1853)]; <i>Pelecyphorus aegrotus limbatus</i> Casey, 1912 [= <i>Philolithus aegrotus aegrotus</i> (LeConte, 1861)]; <i>Pelecyphorus corporalis</i> Casey, 1912, <i>Pelecyphorus reptans</i> Casey, 1912, <i>Pelecyphorus socer</i> Casey, 1912, <i>Pelecyphorus abscissus</i> Casey, 1912, <i>Pelecyphorus fumosus</i> Casey, 1912, <i>Pelecyphorus parvus</i> Casey, 1912, <i>Pelecyphorus morbillosus pacatus</i> Casey, 1912, <i>Pelecyphorus morbillosus sobrius</i> Casey, 1912, <i>Pelecyphorus piceus</i> Casey, 1912, <i>Pelecyphorus piceus crudelis</i> Casey, 1912, <i>Pelecyphorus snowi</i> Casey, 1912, and <i>Pelecyphorus subtenuis</i> Casey, 1912 [= <i>Philolithus morbillosus</i> (LeConte, 1858)]; <i>Bothrasida sanctae-agnae</i> Wilke, 1922 [= <i>Stenomorpha funesta</i> (Champion, 1884)]; <i>Asida flaccida</i> Horn, 1896 [= <i>Stenomorpha embaphionides</i> (Horn, 1894)]; <i>Asida angustula</i> Casey, 1890, <i>Stethasida stricta</i> Casey, 1912, <i>Stethasida muricatula languida</i> Casey, 1912, <i>Stethasida pertinax</i> Casey, 1912, <i>Stethasida socors</i> Casey, 1912, <i>Stethasida angustula inepta</i> Casey, 1912, <i>Stethasida tenax</i> Casey, 1912, and <i>Stethasida vegrandis</i> Casey, 1912 [= <i>Stenomorpha muricatula</i> (LeConte, 1851)]; <i>Stethasida obsoleta expansa</i> Casey, 1912, <i>Stethasida obsoleta opacella</i> Casey, 1912, <i>Stethasida brevipes</i> Casey, 1912, <i>Stethasida torpida</i> Casey, 1912, <i>Stethasida convergens</i> Casey, 1912, <i>Stethasida discreta</i> Casey, 1912, <i>Stethasida longula</i> Casey, 1912, <i>Stethasida adumbrata</i> Casey, 1912, <i>Stethasida occulta</i> Casey, 1912, <i>Stethasida tarsalis</i> Casey, 1912, <i>Stethasida unica</i> Casey, 1912, and <i>Pelecyphorus laevigatus</i> Papp, 1961 [= <i>Stenomorpha obsoleta</i> (LeConte, 1851)]; <i>Trichiasida eremica</i> Wilke, 1922 [= <i>Stenomorpha difficilis</i> (Champion, 1884)]; <i>Trichiasida lineatopilosa</i> Casey, 1912 [= <i>Stenomorpha hirsuta</i> (LeConte, 1851)]; <i>Trichiasida tenella</i> Casey, 1912 [= <i>Stenomorpha hispidula</i> (LeConte, 1851)]; <i>Trichiasida duplex</i> Casey, 1912 [= <i>Stenomorpha villosa</i> (Champion, 1884)]; <i>Alaudes squamosa</i> Blaisdell, 1919, <i>Alaudes testacea</i> Blaisdell, 1919, and <i>Alaudes fallax</i> Fall, 1928 [= <i>Alaudes singularis</i> Horn, 1870]; <i>Edrotes barrowsi</i> Dajoz, 1999 [=<i>Edrotes ventricosus</i> LeConte, 1851]; <i>Nyctoporis tetrica</i> Casey, 1907 and <i>Nyctoporis maura</i> Casey, 1907 [= <i>Nyctoporis aequicollis</i> Eschscholtz, 1831]; <i>Nyctoporis pullata</i> Casey, 1907 [= <i>Nyctoporis sponsa</i> Casey, 1907]; Eleodes tibialis forma oblonga Blaisdell, 1909 [= <i>Eleodes tibialis</i> Blaisdell, 1909]; <i>Eleodes</i> (<i>manni</i> var.) <i>variolosa</i> Blaisdell, 1917 [= <i>Eleodes constrictus</i> LeConte, 1858]; Eleodes cordata forma sublaevis Blaisdell, 1909, Eleodes cordata forma intermedia Blaisdell, 1909, Eleodes cordata forma oblonga Blaisdell, 1909, Eleodes cordata forma elongata Blaisdell, 1909, and <i>Eleodes</i> (<i>cordata</i> var.) <i>adulterina</i> Blaisdell, 1917 [= <i>Eleodes cordata</i> Eschscholtz, 1829]; Eleodes hornii var. monticula Blaisdell, 1918 and <i>Eleodes manni sierra</i> Blaisdell, 1925 [= <i>Eleodes fuchsii</i> Blaisdell, 1909]; Eleodes parvicollis var. squalida Blaisdell, 1918 [= <i>Eleodes parvicollis</i> Eschscholtz, 1829]; <i>Eleodes reflexicollis</i> Mannerheim, 1843 and Eleodes parvicollis forma farallonica Blaisdell, 1909 [= <i>Eleodes planata</i> Eschscholtz, 1829]; <i>Eleodes indentata</i> Blaisdell, 1935 [= <i>Eleodes rotundipennis</i> LeConte, 1857]; <i>Eleodes intricata</i> Mannerheim, 1843 [= <i>Eleodes scabrosa</i> Eschscholtz, 1829]; <i>Eleodes horni fenyesi</i> Blaisdell, 1925 [= <i>Eleodes tenebrosa</i> Horn, 1870]; Eleodes cordata var. horrida Blaisdell, 1918 [= <i>Eleodes tuberculata</i> Eschscholtz, 1829]; <i>Eleodes oblonga</i> Blaisdell, 1933 [= <i>Eleodes versatilis</i> Blaisdell, 1921]; <i>Eleodes dentipes marinae</i> Blaisdell, 1921 [= <i>Eleodes dentipes</i> Eschscholtz, 1829]; Eleodes carbonaria forma glabra Blaisdell, 1909 [= <i>Eleodes carbonaria carbonaria</i> (Say, 1824)]; Eleodes granosa forma fortis Blaisdell, 1909 [= <i>Eleodes granosa</i> LeConte, 1866]; Eleodes pilosa forma ordinata Blaisdell, 1909 [= <i>Eleodes pilosa</i> Horn, 1870]; <i>Trogloderus costatus pappi</i> Kulzer, 1960 [= <i>Trogloderus tuberculatus</i> Blaisdell, 1909]; <i>Trogloderus costatus mayhewi</i> Papp, 1961 [= <i>Trogloderus vandykei</i> La Rivers, 1946]; <i>Bolitophagus cristatus</i> Gosse, 1840 [= <i>Bolitotherus cornutus</i> (Fabricius, 1801)]; <i>Eleates explanatus</i> Casey, 1890 [= <i>Eleates depressus</i> (Randall, 1838)]; <i>Blapstinus sonorae</i> Casey, 1890 [= <i>Blapstinus brevicollis</i> LeConte, 1851]; <i>Blapstinus falli</i> Blaisdell, 1929 [= <i>Blapstinus castaneus</i> Casey, 1890]; <i>Blapstinus brunneus</i> Casey, 1890 and <i>Blapstinus coronadensis</i> Blaisdell, 1892 [=<i>Blapstinus histricus</i> Casey, 1890]; <i>Blapstinus hesperius</i> Casey, 1890 [=<i>Blapstinus intermixtus</i> Casey, 1890]; <i>Blapstinus cinerascens</i> Fall, 1929 [= <i>Blapstinus lecontei</i> Mulsant and Rey, 1859]; <i>Blapstinus niger</i> Casey, 1890 and <i>Blapstinus cribricollis</i> Casey, 1890 [= <i>Blapstinus pimalis</i> Casey, 1885]; <i>Blapstinus arenarius</i> Casey, 1890 [= <i>Blapstinus pratensis</i> LeConte, 1859]; <i>Blapstinus gregalis</i> Casey, 1890 [= <i>Blapstinus substriatus</i> Champion, 1885]; <i>Blapstinus hydropicus</i> Casey, 1890 [= <i>Blapstinus sulcatus</i> LeConte, 1851]; <i>Blapstinus hospes</i> Casey, 1890 [= <i>Blapstinus vestitus</i> LeConte, 1859]; <i>Notibius reflexus</i> Horn, 1894 [= <i>Conibius opacus</i> (LeConte, 1866)]; <i>Notibius affinis</i> Champion, 1885 [=<i>Conibius rugipes</i> (Champion, 1885)]; <i>Conibius parallelus</i> LeConte, 1851 [= <i>Conibius seriatus</i> LeConte, 1851]; <i>Nocibiotes rubripes</i> Casey, 1895 [=<i>Nocibiotes caudatus</i> Casey, 1895]; <i>Nocibiotes gracilis</i> Casey, 1895 and <i>Nocibiotes acutus</i> Casey, 1895 [=<i>Nocibiotes granulatus</i> (LeConte, 1851)]; <i>Conibius alternatus</i> Casey, 1890 [= <i>Tonibius sulcatus</i> (LeConte, 1851)]; <i>Pedinus suturalis</i> Say, 1824 [= <i>Alaetrinus minimus</i> (Palisot de Beauvois