Definition and Reorganization of the Genus Pseudocloeon (Ephemeroptera: Baetidae) With New Species Descriptions and Combinations

Abstract

Labiobaetis Novikova & Kluge (Ephemeroptera: Baetidae) is synonymized with Pseudocloeon Klap?lek on the basis of adult morphology now known from throughout its Holarctic, Afrotropical, Oriental, and Australian range and consistent with that of P. kraepelini Klap?lek, the type of Pseudocloeon. Adult and larval stages of Pseudocloeon are characterized and compared with those of other genera of the Baetis complex. All species previously assigned to Labiobaetis and several species previously assigned to Baetis are reassigned to Pseudocloeon. Tabular summaries of Pseudocloeon species show nomenclatural revisions, regional distribution, and diagnostic character variability among larvae. Pseudocloeon hypodelum, new species, P. inconspicuum, new species, and P. plecttle, new species, are described from Australia. Pseudocloeon involutum, new species, P. petersorum, new species, P. tuberpalpus, new species, P. vitile, new species, P. vultuosum, new species, and P. xeniolum, new species, are described from Papua-New Guinea. Biogeographic data are shown to be congruent with the revised systematics of Pseudocloeon. Occurrences of Pseudocloeon in Papua New Guinea and Australia are probably related to dispersal events as recent as the middle Miocene.