New material and a revision of turtles of the genus Adocus (Adocidae) from the Late Cretaceous of Middle Asia and Kazakhstan

Abstract

This paper reviews shell material of the turtle genus Adocus (Adocidae) from the Late Cretaceous of Middle Asia and Kazakhstan. Three previously recognized species of Adocus from this area are described in detail based on published and new material. The previously recognized species are A. aksary (Dzharakuduk, Uzbekistan; late Turonian), A. foveatus (Kansai, Tajikistan; early Santonian), and A. kizylkumensis (Khodzhakul, Khodzhakulsay and Sheikhdzheili, Uzbekistan; early Cenomanian). Material of additional forms of Adocus are described, two of these are named as new species: Adocus dzhurtasensis, sp. nov. (Dzhurtas, Kazakhstan; Santonian – early Campanian), Adocus bostobensis, sp. nov. (Akkurgan, Baybishe, Buroinak, and Shakh-Shakh, Kazakhstan; Santonian – early Campanian), and Adocus sp. indet. (Itemir, Uzbekistan; Cenomanian).