TWO NEW SPECIES OF TORRENT FROG OF THE GENUS HYLODES (ANURA, HYLODIDAE) WITH NUPTIAL THUMB TUBERCLES

Abstract

We analyzed variation among Hylodes with nuptial tubercles using morphometrics, vocalization, and external morphology. We identified three species: Hylodes phyllodes from the Serra do Mar, State of São Paulo, and southern portion of the State of Rio de Janeiro; a new species from Ilha Grande, municipality of Angra dos Reis; and a new species from the Serra dos Órgãos, municipality of Guapimirim. The new species are both from the state of Rio de Janeiro, southeastern Brazil.