Recent and Tertiary Cocculinidae and Pseudococculinidae (Mollusca: Gastropoda) from New Zealand and New South Wales

Abstract

Abstract The families Cocculinidae and Pseudococculinidae are represented in New Zealand by the following species (fossil taxa asterisked): Cocculinidae — Cocculina cervae Fleming, G. pristina n. sp.*, Coccopigya (nom. nov.) compunctum (Marwick)*, C. otaiana*, C. komitica*, C. hispida, C. crinita, C. oculifera, C. crebrilamina n. spp.; Pseudococculinidae — Pseudococculina gregaria, P. gradata , n. spp.; Notocrater craticulata (Suter), N. maxwelli*, N. gracilis n. spp; Kaiparapelta (n. gen.) singularis n. sp*; Tentaoculus lithodicola, T. neolithodicola, T. haptricola , n. spp; Mesopelex (n. gen.) zelandica n. sp; Kurilabyssia antipodensis n. sp; Caymanabyssia rhina, C. sinespina n. spp; Colotrachelus (n. gen.) hestica n. sp. New South Wales representatives are the cocculinids Coccopigya crinita and C. barbatula n. spp., and the pseudococculinids Pseudococculina gregaria, Notocrater ponderi , and Kurilabyssia antipodensis n. spp. A new subfamily, Caymanabyssiinae, is proposed for Caymanabyssia Moskalev and Colotrachelus n. gen. Tentaoculus lithodicola and T. neolithodicola live on the carapaces of living deep‐sea stone crabs (Lithodidae), while T. haptricola eats decaying algal holdfasts at bathyal depths; unique and hitherto unrecorded associations for the Mollusca. So far as is known, all other species recorded use waterlogged wood as a substratum and food source. Cocculinid and pseudococculinid phylogeny and distribution are discussed. Cocculinella coercita (Hedley) (Cocculinellidae) is re‐illustrated and compared with C. kopua and C. osteophila Marshall. Lectotypes are designated for Pseudococculina rugosoplicata Schepman and Acmaea parva var. tasmanica Pilsbry. Cocculina meridionalis (Hedley) and Acmaea eccentrica Test are synoymised with Acmaea parva var. tasmanica Pilsbry, which together with Scutellina alboradiata Verco, is referred to Propilidium Forbes and Hanley (Lepetidae).